Professor Titular da Universidad Nacional Autónoma de México (UNAM). Possui graduação em Ciencias Biológicas - Universidad de Costa Rica (1984), mestrado em Ecologia - Pennsylvania State University (1990) e doutorado em Ecologia - Pennsylvania State University (1993).
bachelor's at Ciencias Biológicas from Universidad de Costa Rica (1984), master's at Ecologia from Pennsylvania State University (1990) and doctorate at Ecologia from Pennsylvania State University (1993).